The engrailed and small engrailed (''Ectropis crepuscularia'') are

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

s of the family

Geometridae The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek ''geo'' γεω (derivative form of or "the earth"), and ''met ...

found from the

British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...

through central and eastern Europe to the

Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admini ...

and

Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...

. The western

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...

and

Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...

and the

Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...

represent the southern limit of the distribution (with the

Balkan countries The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the whol ...

). In the north, the distribution area ends at the Arctic Circle. It also occurs in North America. Debate exists as to whether they make up one

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

, or whether ''E. crepuscularia'' actually refers only to the small engrailed, with the engrailed proper being separable as ''E. bistortata''. The ground colour of the wings is buff or grey, variably marked with darker

fascia A fascia (; plural fasciae or fascias; adjective fascial; from Latin: "band") is a band or sheet of connective tissue, primarily collagen, beneath the skin that attaches to, stabilizes, encloses, and separates muscles and other internal organs. ...

and a pale postdiscal crossline. The darker markings are not usually as strong as in the rather similar willow beauty. Melanic forms occur fairly frequently. The

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...

is . One or two broods are produced each year. In the British Isles, the adults can be seen at any time between March and August; this time range may vary in other parts of this moth's range. The species flies at night and is attracted to light. The greyish

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...

is truly

polyphagous Feeding is the process by which organisms, typically animals, obtain food. Terminology often uses either the suffixes -vore, -vory, or -vorous from Latin ''vorare'', meaning "to devour", or -phage, -phagy, or -phagous from Greek φαγε ...

, feeding on a huge range of

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...

s. As a caterpillar, the species is known as the saddleback looper. The species overwinters as a

pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...

Recorded food plants

* ''Acer'',

maple ''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since http ...

* ''

Aconitum ''Aconitum'' (), also known as aconite, monkshood, wolf's-bane, leopard's bane, mousebane, women's bane, devil's helmet, queen of poisons, or blue rocket, is a genus of over 250 species of flowering plants belonging to the family Ranunculaceae. ...

'', monkshood * ''Alnus'',

alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...

* ''

Aquilegia ''Aquilegia'' (common names: granny's bonnet, columbine) is a genus of about 60–70 species of perennial plants that are found in meadows, woodlands, and at higher altitudes throughout the Northern Hemisphere, known for the spurred petals Pu ...

'', columbine * ''Betula'',

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...

* ''

Calluna vulgaris ''Calluna vulgaris'', common heather, ling, or simply heather, is the sole species in the genus ''Calluna'' in the flowering plant family Ericaceae. It is a low-growing evergreen shrub growing to tall, or rarely to and taller, and is found wid ...

'', heather * ''

Camellia japonica ''Camellia japonica'', known as common camellia, or Japanese camellia, is a species of flowering plant in the family Theaceae. There are thousands of cultivars of ''C. japonica'' in cultivation, with many colors and forms of flowers. In the U.S. ...

'', Japanese camellia * ''Castanea'',

chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrelat ...

* ''Centaurea'',

star thistle ''Centaurea'' () is a genus of over 700 species of herbaceous thistle-like flowering plants in the family Asteraceae. Members of the genus are found only north of the equator, mostly in the Eastern Hemisphere; the Middle East and surrounding reg ...

* ''

Cirsium arvense ''Cirsium arvense'' is a perennial species of flowering plant in the family Asteraceae, native throughout Europe and western Asia, northern Africa and widely introduced elsewhere.Joint Nature Conservation Committee''Cirsium arvense'' The standa ...

'', creeping thistle * ''Cornus'',

dogwood ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous trees or shrub ...

* ''

Daphniphyllum ''Daphniphyllum'' is the sole genus in the flowering plant family Daphniphyllaceae and was described as a genus in 1826. The genus includes evergreen shrubs and trees mainly native to east and southeast Asia, but also found in the Indian Subconti ...

'' * ''

Diervilla ''Diervilla'', or bush honeysuckle, is a genus of three species of deciduous shrubs in the family Caprifoliaceae, all indigenous to eastern North America. The genus is named after a French surgeon Dr. Marin Diereville, who introduced the plant ...

'', bush honeysuckle * ''Frangula'',

alder buckthorn ''Frangula alnus'', commonly known as alder buckthorn, glossy buckthorn, or breaking buckthorn, is a tall deciduous shrub in the family Rhamnaceae. Unlike other "buckthorns", alder buckthorn does not have thorns. It is native to Europe, northern ...

* ''Fraxinus'',

ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non- gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...

* ''Genista'',

broom A broom (also known in some forms as a broomstick) is a cleaning tool consisting of usually stiff fibers (often made of materials such as plastic, hair, or corn husks) attached to, and roughly parallel to, a cylindrical handle, the broomstick. I ...

* ''

Glycine Glycine (symbol Gly or G; ) is an amino acid that has a single hydrogen atom as its side chain. It is the simplest stable amino acid (carbamic acid is unstable), with the chemical formula NH2‐ CH2‐ COOH. Glycine is one of the proteinogeni ...

'', soybean * ''

Hypericum maculatum ''Hypericum maculatum'', commonly known as imperforate St John's-wort, or spotted St. Johnswort, is a species of perennial herbaceous flowering plant in the family Hypericaceae. It is native to Europe and Western Asia where it grows in moist m ...

'', imperforate St. John's wort * ''Ilex'',

holly ''Ilex'' (), or holly, is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen o ...

* ''Juglans'',

walnut A walnut is the edible seed of a drupe of any tree of the genus ''Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. Although culinarily considered a "nut" and used as such, it is not a true ...

* ''Larix'',

Larch Larches are deciduous conifers in the genus ''Larix'', of the family Pinaceae (subfamily Laricoideae). Growing from tall, they are native to much of the cooler temperate northern hemisphere, on lowlands in the north and high on mountains furt ...

* ''

Lindera Dried fruits of ''Lindera neesiana'' used as spice (coll. MHNT) ''Lindera'' is a genus of about 80–100honeysuckle Honeysuckles are arching shrubs or twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae, native to northern latitudes in North America and Eurasia. Approximately 180 species of honeysuckle have been identified in both contin ...

* ''Lythrum salicaria'',

purple loosestrife ''Lythrum salicaria'' or purple loosestrifeFlora of NW Europe''Lythrum salicaria'' is a flowering plant belonging to the family Lythraceae. It should not be confused with other plants sharing the name loosestrife that are members of the family P ...

* ''

Malus ''Malus'' ( or ) is a genus of about 30–55 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples, wild apples, and rainberries. The genus is native to the temperate zone of th ...

'', apple * ''

Philadelphus ''Philadelphus'' () (mock-orange) is a genus of about 60 species of shrubs from 3–20 ft (1–6 m) tall, native to North America, Central America, Asia and (locally) in southeast Europe. They are named "mock-orange" in reference to their ...

'', mock-orange * ''Picea'',

spruce A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...

* '' Pieris'' * ''Pinus'',

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...

* ''

Plectranthus ''Plectranthus'' is a genus of about 85 species of flowering plants from the sage family, Lamiaceae, found mostly in southern and tropical Africa and Madagascar. Common names include spur-flower. ''Plectranthus'' species are herbaceous perennia ...

'' * ''

Populus ''Populus'' is a genus of 25–30 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar (), aspen, and cottonwood. The we ...

'', poplar * ''Pseudotsuga'',

Douglas fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three va ...

* ''Quercus'',

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

* ''Ribes rubrum'',

redcurrant The redcurrant or red currant (''Ribes rubrum'') is a member of the genus ''Ribes'' in the gooseberry family. It is native to western Europe. The species is widely cultivated and has escaped into the wild in many regions. Description ''Ribes r ...

* ''Rosa'',

rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...

* ''Rubus idaeus'',

raspberry The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with w ...

Hübner (1825) ''Ectropis''
/ref> * ''

Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distribu ...

'', dock * ''Salix'',

willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...

* ''Sambucus'', elder * ''Shepherdia canadensis'',

Canada buffaloberry ''Shepherdia canadensis'', commonly called Canada buffaloberry, russet buffaloberry, soopolallie, soapberry, or foamberry (Ktunaxa: ',) is one of a small number of shrubs of the genus ''Shepherdia'' that bears edible berries. Description The pla ...

* ''Sorbus'' spp.,

rowan The rowans ( or ) or mountain-ashes are shrubs or trees in the genus ''Sorbus ''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' (''s.l.'') are commonly known as whitebeam, r ...

s * ''Spartina'',

cordgrass ''Spartina'' is a taxon of plants in the grass family, frequently found in coastal salt marshes. Its species are commonly known as cordgrass or cord-grass, and are native to the coasts of the Atlantic Ocean in western and southern Europe, north ...

* ''

Thuja ''Thuja'' ( ) is a genus of coniferous tree or shrub in the Cupressaceae (cypress family). There are five species in the genus, two native to North America and three native to eastern Asia. The genus is monophyletic and sister to ''Thujopsis''. M ...

'' * ''Trifolium'',

clover Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus ...

* ''

Tsuga ''Tsuga'' (, from Japanese (), the name of ''Tsuga sieboldii'') is a genus of conifers in the subfamily Abietoideae of Pinaceae, the pine family. The common name hemlock is derived from a perceived similarity in the smell of its crushed folia ...

'', hemlock * ''Ulmus'',

elm Elms are deciduous and semi-deciduous trees comprising the flowering plant genus ''Ulmus'' in the plant family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of North ...

* ''

Vaccinium ''Vaccinium'' is a common and widespread genus of shrubs or dwarf shrubs in the heath family (Ericaceae). The fruits of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry (whort ...

'' * ''

Zanthoxylum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae in ...

References

External links

Engrailed on ''UKMoths''
*
''Lepiforum e.V.''
Boarmiini Moths described in 1775 Moths of Asia Moths of Europe Moths of North America Taxa named by Michael Denis Taxa named by Ignaz Schiffermüller {{Boarmiini-stub

Recorded food plants

References

Further reading

External links